Scientists & Staff
Alexander L. Friend
Office of the Director
Albany, CA 94710
As the Director of the Pacific Southwest Research Station (PSW), Dr. Friend leads the efforts of approximately 50 agency scientists working in California, Hawaii and the U.S.-affiliated Pacific Islands. He came to PSW from the agency's Washington D.C. headquarters, where he served as Budget Coordinator responsible for planning, formulating, monitoring, and executing the USDA Forest Service’s $300 million Research and Development budget. During his four-year assignment, Dr. Friend worked with Station Directors nationwide to foster a better understanding of and involvement in the budget process; served as a liaison with key stakeholders; and improved budget operations within the agency.
Dr. Friend joined the Forest Service in 2002 as a project leader and research ecologist, leading the Ecosystems unit at the then-North Central Research Station in Houghton, Michigan. He also assisted with the merger of the Station with Northeastern Research Station to form Northern Research Station.
Before joining the Forest Service, Dr. Friend was a professor of forestry at Mississippi State University from 1989 to 2001. He taught both graduate and undergraduate courses in forest biology.
- University of Washington, Ph.D. ,
- North Carolina State University, M.S. ,
- The University of the South., B.S. ,
- Ecological Society of America
- Society of American Foresters
- Soil Science Society of America
Publications & Products
- Kodrzycki, R.J.; Michaels, R.B.; Friend, A.L.; Zalesny, R.S., Jr..; Mawata, Ch.P.; McDonald, D.W. . 2008. Non-destructive digital imaging in poplar allows detailed analysis of adventitious rooting dynamics. In: 5th international symposium on adventitious root formation: from cell fate flexibility to root meristem determination and biomass formation; 2008 June 16-20; Madrid, Spain. [Place of publication unknown]; [Publisher name unknown]: 33-34.
- King, J.S.; Giardina, C.P.; Pregitzer, K.S.; Friend, A.L. . 2007. Biomass partitioning in red pine (Pinus resinosa) along a chronosequence in the Upper Peninsula of Michigan. Canadian Journal of Forest Research. 37: 93-102.
- Zalesny, Ronald S., Jr.; Friend, A. L.; Kodrzycki, B.; McDonald, D.W.; Michaels, R.; Wiese, A.H.; Powers, J.W. . 2007. Using low energy x-ray radiography to evaluate root initiation and growth of Populus. In: Annual meeting of the poplar council of Canada: Poplar culture: a collaborative effort from clone to mill; 2007 September 16-21; Quebec City, Quebec, Canada. [Place of publication unknown]:[Publisher unknown]:73.
- Roberts, Scott D.; Friend, Alex L.; Schoenholtz, Stephen H. . 2006. Growth of precommercially thinned loblolly pine 4 years following application of poultry litter. Gen. Tech. Rep. SRS-92. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. pp. 139-142
- Kubiske, Mark E.; Quinn, Vanessa A.; Heilman, Warren E.; McDonald, Evan P.; Marquardt, Paula E.; Teclaw, Ron M.; Friend, Alexander L.; Karnosky, David F. . 2006. Interannual climatic variation mediates elevated CO2 and O3 effects on forest growth. Global Change Biology. 12: 1054-1068.
- Nowak, Jaroslaw; Friend, Alexander L. . 2006. Loblolly pine and slash pine responses to acute aluminum and acid exposures. Tree Physiology 26, 1207-1215.
- Friend, Alexander L.; Roberts, Scott D.; Schoenholtz, Stephen H.; Mobley, Juanita A.; Gerard, Patrick D. . 2006. Poultry litter application to loblolly pine forests: growth and nutrient containment. Published in J. Environ. Qual. 35:837-848 (2006).
- Nowak, Jaroslaw; Friend, Alexander L. . 2005. Aluminum fractions in root tips of slash pine and loblolly pine families differing in Al resistance. Tree Physiology 25, 245-250
- Karberg, N.J.; Pregitzer, K.S.; King, J.S.; Friend, A.L.; Wood, J.R. . 2005. Soil carbon dioxide partial pressure and dissolved inorganic carbonate chemistry under elevated carbon dioxide and ozone. Oecologia. 142: 296-306.
- Coleman, Mark D.; Friend, Alexander L.; Kern, Christel C. . 2004. Carbon allocation and nitrogen acquisition in a developing Populus deltoides plantation. Tree Physiology 24, 1347-1357
- Coleman, Mark D.; Friend, Alexander L.; Kern, Christel C. . 2004. Carbon allocation and nitrogen acquisition in a developing Populus deltoides plantation. Tree Physiology 24:1347-1357
- Kern, Christel C.; Friend, Alexander L.; Johnson, Jane M.; Coleman, Mark D. . 2004. Fine root dynamics in a developing Populus deltoides plantation. Tree Physiology. Volume 24, 2004. pp. 651-660.
- Kern, Christel C.; Friend, Alexander L.; Johnson, Jane M.-F.; Coleman, Mark D. . 2004. Fine root dynamics in a developing Populus deltoides plantation. Tree Physiology 24: 651-660
- Karberg, N.J.; Pregitzer, K.S.; King, J.S.; Friend, A.L.; Wood, J.R. . 2004. Soil carbon dioxide partial pressure and dissolved inorganic carbonate chemistry under elevated carbon dioxide and ozone. Oecologia (2005) 142: 296-306
- Roberts, Scott D.; Friend, Alexander L.; Gerard, Patrick D. . 2004. The Effect of Large Applications of Nutrients From Organic Waste on Biomass Allocation and Allometric Relations in Loblolly Pine. Gen. Tech. Rep. SRS–71. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. pp. 398-402
- Woolfolk, Walter T.M.; Friend, Alexander L. . 2003. Growth response of cottonwood roots to varied NH4:NO3 ratios in enriched patches. Tree Physiology (2003) 23, 427-432
- Kubisiak, Thomas L.; Nelson, C. Dana; Nowak, J.; Friend, A.L. . 2000. Genetic Linkage Mapping of Genomic Regions Conferring Tolerance to High Aluminum in Slash Pine. Journal of Sustainable Forestry, Vol. 10, No. 1/2, 2000, pp. 69-78
- Friend, Alexander L.; Coleman, Mark D.; Isebrands, J.G. . 1994. Carbon allocation to root and shoot systems of woody plants. In: Biology of Adventitious Root Formotion, Edited by T.D. Davis and B.E. Haissig. Plenum Press. New York. 1994 p. 245-274